Publications

Research papers

2024. Pyrcz, T. W., Boyer, P., Petit, J.-C., Garlacz, R., Zając-Garlacz, K., Espeland, M., Willmott, K. R. Bedazzled: a new, striking species of Corades from the outskirts of Quito questions our knowledge of Andean cloud forest butterflies (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 5453(2): 255-262. DOI: 10.11646/zootaxa.5453.2.6 (online 20 May 2024).

2024. Freitas, A. V. L., Braga, L., Rosa, A. H. B., Willmott, K. R. Rediscovery of Adelpha herbita herbita Weymer more than a century after its description (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 34(1): 40-43. DOI: 10.5281/zenodo.10882153 (online 5 April 2024).

2024. Carvalho, S., A. P., Owens, H. L., St Laurent, R. A., Earl, C., Dexter, K. M., Messcher, R. L., Willmott, K. R., Aduse-Poku, K., Collins, S. C., Homziak, N. T., Hoshizaki, S., Hsu, Y.-F., Kizhakke, A. G., Kunte, K., Martins, D. J., Mega, N. O., Morinaka, S., Peggie, D., Romanowski, H. P., Sáfián, S., Vila, R., Wang, H., Braby, M. F., Espeland, M., Breinhold, J., Pierce, N. E., Kawahara, A. Y., Lohman, D. J. Comprehensive phylogeny of Pieridae butterflies reveals strong correlation between diversification and temperature. iScience 27: 109336, 1-13. DOI: 10.1016/j.isci.2024.109336 (online 28 February 2024).

2023. Willmott, K. R., Lamas, G., Hall, J. P. W., Boyer, P., Pyrcz, T., Florcyzk, K. 2023. A new species and eight new subspecies of high elevation Actinote from Colombia, Ecuador and Peru (Nymphalidae, Heliconiinae, Acraeini). Tropical Lepidoptera Research 34(1): 1-20. DOI: 10.5281/zenodo.10027609 (online 27 October 2023).

2023. Garzón-Orduña, I. J., Silva-Brandão, K. L., Willmott, K. R., Freitas, A. V. L., Wahlberg, N., Brower, A. V. Z. 2023. Wing pattern diversity in Eunica butterflies (Nymphalidae: Biblidinae): phylogenetic analysis reveals decoupled adaptive trends in dorsal sexual dimorphism and ventral eyespot evolution. Cladistics 2023: 1-20. DOI: 10.1111/cla.12556 (online 15 September 2023).

2023. Pyrcz, T. W., Fahråeus, C., Boyer, P., Mahecha, O., Lorenc-Brudecka, J., Zając, K. S., Willmott, K. R., Padrón, S. The sunny butterflies: new species of high montane pierids of the Catasticta poujadei group from the Peruvian Andes (Pieridae, Pierinae, Aporiina). Zootaxa 5336(4): 530-542. DOI: 10.11646/zootaxa.5336.4.4 (online 24 August 2023).

2023. Rush, C. E., Freitas, A. V. L., Magaldi, L., Willmott, K. R., Hill, R. I. One generalist or several specialists? Comparative analysis of the polyphagous butterfly Adelpha serpa celerio and its serpa-group relatives. Zoologica Scripta 52(5): 475-493. DOI: 10.1111/zsc.12615 (online 22 June 2023).

2023. Nakahara, S., Kleckner, K., Barbosa, E. P., Lourenço, G. M., Casagrande, M. M., Willmott, K. R., Freitas, A. V. L. Reassessment of the type locality of Euptychia stigmatica Godman, 1905, with the descriptions of two new sibling species from Amazonia (Lepidoptera, Nymphalidae, Satyrinae, Satyrini). ZooKeys 1167: 57-88. DOI: 10.389/zookeys.1167.102979 (online 14 June 2023).

2023. Kawahara, A. Y., …. Willmott, K. R., … A global phylogeny of butterflies reveals their evolutionary history, ancestral hosts, and biogeographic origins. Nature Ecology & Evolution DOI: 10.1038/s41559-023-02041-9 (online 15 May 2023).

2023. Mota, L. L., Santos, J. P., Willmott, K. R., Freitas, A. V. L. Butterfly assemblages differ among vegetation types in Southern Amazonia. Diversity 2023, 15: 624. DOI: 10.3390/d15050624 (online 4 May 2023).

2023. Espeland, M., Nakahara, S., Zacca, T., Barbosa, E. P., Huertas, B.,  Marín, M. A., Lamas, G., Benmesbah, M., Brévignon, C., Casagrande, M. M., Fahråeus, C., Grishin, N., Kawahara, A. Y., Mielke, O. H. H., Miller, J. Y., Nakamura, I., Navas, V., Patrusky, B., Pyrcz, T. W., Richards, L., Tan, D., Tyler, S., Viloria, A., Warren, A. D., Xiao, L., Freitas, A. V. L., Willmott, K. R. Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Satyrinae, Nymphalidae). Systematic Entomology 2023; 1-73 DOI: 10.1111/syen/12590 (online 10 April 2023).

2023. Hall, J. P. W., Willmott, K. R., Ahrenholz, D. H. A review of the Argyrogrammana amalfreda group fauna of the western Amazon (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(Supplement 1): 61-85. DOI: 10.5281/zenodo.7799089 (online 7 April 2023).

2023. Hall, J. P. W., Willmott, K. R., Ahrenholz, D. H. A review of the Argyrogrammana fauna of the eastern Andes (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(Supplement 1): 49-60. DOI: 10.5281/zenodo.7799076 (online 7 April 2023).

2023. Hall, J. P. W., Willmott, K. R. A review of the Argyrogrammana fauna of the Transandean region (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(Supplement 1): 28-48. DOI: 10.5281/zenodo.7799056 (online 7 April 2023).

2023. Doré, M., Willmott, K. R., Lavergne, S., Chazot, N., Freitas, A. V. L., Fontaine, C., Elias, M. Mutualistic interactions shape global spatial congruence and climatic niche evolution in Neotropical mimetic butterflies. Ecology Letters 26: 843-857. DOI: 10.1111/ele.14198 (online 17 March 2023).

2022. Dietz, L., Eberle, J., Mayer, C., Kukowka, S., Bohacz, C., Baur, H., Espeland, M., Huber, B. A., Hutter, C., Mengual, X., Peters, R. S., Vences, M., Wesener, T., Willmott, K. R., Misof, B., Niehuis, O., Ahrens, D. Standardized nuclear markers advance metazoan taxonomy. Methods in Ecology and Evolution, 14(2): 543-555. DOI: 10.1111/2041-210X.14041 (online 12 December 2022).

2022. Pyrcz, T. Willmott, K. R., Lamas, G., Boyer, P., Florczyk, K. Fahråeus, C., Mahecha, O., Cerdeña, J., Mrozek, A., Farfán, J., Zubek, A. Considerations on the systematics of Neotropical Pierina, with the description of two new species of Phulia Herrich-Schäffer from the Peruvian Andes. Neotropical Entomology, DOI: 10.1007/s13744-022-00999-y (online 16 November 2022).

2022. Callaghan, C. T., … Willmott, K. R., … The benefits of contributing to the citizen science platform iNaturalist as an identifier. PLoS Biology 20(11): e3001843. DOI: 10.1371/journal.pbio.3001843 (online 10 November 2022).

2022. Mota, L., Boddington, S. J., Brown, K. S., Callaghan, C. J., Carter, G., Carter, W., Dantas, S. M., Dolibaina, D. R., Garwood, K., Hoyer, R. C., Robbins, R. K., Soh, A., Willmott, K. R., Freitas, A. V. L. The butterflies of Cristalino Lodge, in the Brazilian southern Amazonia: An updated species list with a significant contribution from citizen science. Biota Neotropica, 22(3): e20221367, 2022, 1-25. DOI: 10.1590/1676-0611-BN-2022-1367 (online 7 November 2022).

2022. Pyrcz, T. W., Lachowska-Cierlik, D., Willmott, K. R., Mrozek, A., Machecha-Jiménez, O., Fåhraeus, C., Boyer, P., Martín, S., Espeland, M. A new genus in the diverse Andean Pedaliodes complex uncovered using target enrichment (Lepidoptera, Nymphalidae). Systematic Entomology, 2022; 1-15. DOI: 10.1111/syen.12568 (online 1 October 2022).

2022. Nakahara, S., Huertas, B., Nandi Das, G., Willmott, K. R. Revision of the “celia clade” of Pseudodebis Forster, 1964, with two new species and notes on Papilio phorcys Fabricius, 1793 (Lepidoptera: Nymphalidae: Satyrinae). Neotropical Entomology, 51: 536-556. DOI: 10.1007/s13744-022-00954-x (online 8 June 2022).

2022. Mota, L. L., Rosa, A. H. B., Vasconcellos, L. R., Willmott, K. R., Freitas, A. V. L. A new subspecies of Mechanitis lysimnia from southern Amazonia (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research, 32(1): 47-51. DOI: 10.5281/zenodo.6604683 (online 1 June 2022).

2022. Checa, M. F., Nogales, S., Salazar, P. A., Bustos, L., Ojeda, V., Bustos, A., Willmott, K. R. Implementing a novel approach to long-term monitoring of butterfly communities in the Neotropics. Insect Conservation and Diversity, 2022: 1-13. DOI: 10.1111/icad.12567 (online 2 February 2022). Online stories: Florida Museum of Natural History, Atmos, Mongabay.

2022. Nakahara, S., Rodríguez-Melgarejo, M., Kleckner, K., Espeland, M., Casagrande, M. M., See, J., Gallice, G., Corahua, T., Barbosa, E. P., Lamas, G., Willmott, K. R. Systematic revision of a new butterfly genus, Cisandina Nakahara & Espeland, n. gen., with descriptions of three new taxa (Lepidoptera: Nymphalidae: Satyrinae). Insect Systematics and Diversity, 6(1): 2; 1-30. DOI: 10.1093/isd/ixab028 (online 6 January 2022).

2021. Nakahara, S., Willmott, K. R., MacDonald, J. R., Thurman, A. Description of an enigmatic new species of Chloreuptychia Forster, 1964 from Panama (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research, 31(3): 145-150. DOI: 10.5281/zenodo.5776918 (online 17 December 2021).

2021. Doré, M., Willmott, K. R., Leroy, B., Chazot, N., Mallet, J., Freitas, A. V. L., Hall, J. P. W., Lamas, G., Dasmahapatra, K. K., Fontaine, C., Elias, M. Anthropogenic pressures coincide with Neotropical biodiversity hotspots in a flagship butterfly group. Diversity and Distributions 1-19. DOI: 10.1111/ddi.13455 (online 9 December 2021).

2021. Mahecha-J., O., Florczyk, K., Willmott, K. R., Cerdeña, J., Zubek, A., Boyer, P., Farfán, J., Lachowska-Cierlik, D., Pyrcz, T. W. Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru: description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina). Zootaxa 5072(3): 201-237. DOI: 10.11646/zootaxa.5072.3.1 (online 30 November 2021).

2021. Rosa, A. H. B., Huertas, B., Willmott, K. R., Barbosa, E. P., Machado, P. A., Mielke, O. H. H., Canaan, C. H. P., Freitas, A. V. L. Fifty years without a name: a new species of Splendeuptychia Forster (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5061(1): 95-114. DOI: 10.11646/zootaxa.5061.1.4 (online 3 November 2021).

2021. Núñez, R., Willmott, K. R., Álvarez, Y., Genaro, J., Pérez-Asso, A., Quejereta, M., Turner, T., Miller, J. Y., Brévignon, C., Lamas, G., Hausmann, A. Integrative taxonomy clarifies species limits in the hitherto monotypic passion-vine butterfly genera Agraulis and Dryas (Lepidoptera, Nymphalidae, Heliconiinae). Systematic Entomology, 47: 152-178. DOI: 10.1111/syen/12523 (online 16 October 2021; in issue 11 Jan 2022).

2021. Willmott, K. R., Lamas, G., Hall, J. P. W., Vitale, F., Boyer, P., Petit, J.-C., Radford, J., Elias, M. A new species and thirty-eight new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research, 31(Supplement 3): 1-80. DOI: 10.5281/zenodo.5526837 (online 30 September 2021).

2021. Zacca, T., Casagrande, M. M., Mielke, O. H. H., Huertas, B., Freitas, A. V. L., Marín, M. A., Espeland, M., Willmott, K. R. A new euptychiine butterfly species from south Brazil and taxonomic rearrangements for Taydebis Freitas, 2013 and Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5023(4): 555-570. DOI: 10.11646/zootaxa.5023.4.5 (published 18 August 2021)

2021. Padrón, P. S., Pyrcz, T. W., Willmott, K. R. A contribution towards resolving the systematics of the high altitude tropical Andean satyrine genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae [sic]: Satyrinae). Neotropical Entomology, 37 pp. DOI: 10.1007/s13744-021-00897-9 (online 4 August 2021).

2021. Nakahara, S., Matos-Maraví, P., Willmott, K. R., Nakamura, I., MacDonald, J. R. Description of a new species of Pseudodebis Forster, 1964 from Central America (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research, 31(1): 35-41. DOI: 10.5281/zenodo.4721627 (online 3 May 2021).

2021. Tan, D., Parus, A., Dunbar, M., Espeland, M., Willmott, K. R. COI barcode species delineation methods imply critically underestimated diversity in “common” Hermeuptychia butterflies (Lepidoptera: Nymphalidae: Satyrinae). Zoological Journal of the Linnean Society, 193(4): 1256-1270. DOI: 10.1093/zoolinnean/zlab007 (online 10 March 2021).

2021. Paéz, V., E., Valkonen, J. K., Willmott, K. R., Matos-Maraví, P., Elias, M., Mappes, J. Hard to catch: Experimental evidence supports evasive mimicry. Proceedings of the Royal Society of London B, 288: 20203052; 1-10. DOI: 10.1098/rspb.2020.3052 (online 10 March 2021). Story in Treehugger; Sciences et Avenir; CNRS.

2021. Nakahara, S., Matos-Maraví, P., Schwartz, J., Willmott, K. R. Assessing generic synapomorphy of Pseudodebis Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae) and a recent speciation between two new species with a shift in elevation in the western Andes. Invertebrate Systematics, 35: 158-180. DOI: 10.1071/IS20024 (online 8 Feb 2021).

2020. Nakahara, S., Lamas, G., Willmott, K. R., Espeland, M. Description of a new genus and species for a common and widespread Amazonian satyrine butterfly (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). PeerJ, 8:e10324, 1-25. DOI: 10.7717/peerj.10324. (online 4 December 2020).

2020. Nakahara, S., Kleckner, K., Lamas, G., Huertas, B., Willmott, K. R. A contribution towards the systematics of Magneuptychia Forster, 1964: Caeruleuptychia francisca (Butler, 1870), n. comb. (Lepidoptera: Nymphalidae: Satyrinae). Tijdschrift voor Entomologie, 163: 51-61. DOI: 10.1163/22119434-bja10004 (online 20 October 2020).

2020. Ortiz-Acevedo, E., Gomez, J., Willmott, K. R., Espeland, M., Toussaint, E. The roles of wing color pattern and geography in the evolution of Neotropical Preponini butterflies. Ecology and Evolution, 10: 12801-12816. DOI: 10.1002/ece3.6816 (online 3 Oct 2020).

2020. Zacca, T., Casagrande, M. M., Mielke, O H. H., Huertas, B., Espeland, M., Freitas, A. V. L., Willmott, K. R., Nakahara, S., Lamas, G. Revalidation of Vareuptychia Forster, 1964, description of Vanima, gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 4858(1): 001-034. DOI: 10.11646/zootaxa.4858.1.1 (online 28 September 2020).

2020. Zacca, T., Casagrande, M., Mielke, O. H. H., Huertas, B., Barbosa, E., Freitas, A. V. L., Willmott, K. R. Description of Emeryus gen. nov. to accommodate three species formerly placed in Paryphthimoides Forster, 1964 (Lepidoptera: Nymphalidae, Satyrinae). Austral Entomology 59: 505-523. DOI: 10.1111/aen.12479 (online 1 Sep 2020).

2020. Willmott, K. R., Lamas, G., Hall, J. P. W., Mota, L. L., Kell, T. The common, the rare, and the lost: Descriptions of twelve new species and three new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research, 30(Supplement 1): 1-49. DOI: 10.5281/zenodo.3990663 (online 24 Aug 2020).

2020. Padrón, P. S., Vélez, A., Miorelli, N., Willmott, K. R. Urban areas as refuges for endemic fauna: description of the immature stages of Catasticta flisa duna (Eitschberger & T. Racheli, 1998) (Lepidoptera: Pieridae) and its ecological interactions. Neotropical Biodiversity 6(1): 109-116. DOI 10.1080/23766808.2020.1769993 (online 19 May 2020).

2020. Zacca, T., Casagrande, M. M., Mielke, O. H. H., Huertas, B., Barbosa, E. P., Freitas, A. V. L., Lamas, G., Espeland, M., Brévignon, C., Nakahara, S., Checa, M. F., Willmott, K. R. Systematics of the Neotropical butterfly genus Paryphthimoides Forster, 1964 (Lepidoptera: Nymphalida: Satyrinae), with descriptions of seven new taxa. Insect Systematics and Evolution DOI 10.1163/1876312X-00001027, 1-55 (online 14 Feb 2020).

2019. Nakahara, S., Barbosa, E. P., Willmott, K. R., Lamas, G., Freitas, A. V. L. Two new species of Taygetina Forster, 1964 from Ecuador, Peru and Brazil with a possible case of “juxta loss” in butterflies (Lepidoptera: Nymphalidae: Satyrinae). Insect Systematics and Diversity 3(6): 9; 1-13. DOI: 10.1093/isd/ixz023 (online 20 Nov 2019).

2019. Nakahara, S., Zacca, T., Dias, F. M. S., Dolibaina, D. R., Xiao, L., Espeland, M., Casagrande, M. M., Mielke, O. H. H., Lamas, G., Huertas, B., Kleckner, K., Willmott, K. R. Revision of the poorly known Neotropical butterfly genus Zischkaia Forster, 1964 (Lepidoptera, Nymphalidae, Satyrinae), with descriptions of nine new species. European Journal of Taxonomy 551: 1-67. DOI: 10.5852/ejt.2019.551 (online 12 Sep 2019).

2019. Brévignon, C., Rosant, T., Lamas, G., Tyler, S., Willmott, K. R. Description d’une nouvelle espèce d’Euptychiina du genre Chloreuptychia (Lepidoptera, Nymphalidae, Satyrinae). Bulletin de la Société Entomologique de France 124(2): 127-138. DOI: 10.32475/bsef_2044 (June 2019).

2019. Freitas, A. V. L., Woodbury, M. S., Barbosa, E. P., Willmott, K. R. A new subspecies of Adelpha messana (C. Felder & R. Felder) from the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 29(2): 62-66. DOI: 10.5281/zenodo.2685490 (online 17 May 2019).

2019. Chazot, N., Willmott, K. R., Lamas, G., Freitas, A., Piron-Prunier, F., Arias, C., Mallet, J., De Silva, D. L., Elias, M. Renewed diversification following Miocene landscape turnover in a Neotropical butterfly radiation. Global Ecology and Biogeography 2019; 1-15. DOI: 10.1111/geb.12919 (online 10 May 2019).

2019. Willmott, K. R., Marín, M. A., Nakahara, S., Pomerantz, T., Lamas, G., Espeland, M., Xiao, L., Hall, J. P. W., Robinson Willmott, J. I., Freitas, A. V. L. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research 29(1): 29-44 (online 3 May 2019).

2019. Torres, K. P., Artieda, N., Salazar, P., Willmott, K. R., Hill, R. I. 2019. Life history descriptions of Adelpha attica attica, Adelpha epione agilla, and Adelpha jordani from an eastern Ecuador lowland forest. Tropical Lepidoptera Research 29(1): 19-28 (online 3 May 2019).

2019. Nakahara, S., Lamas, G., Tyler, S., Marín, M., Huertas, B., Willmott, K. R., Mielke, O. H. H., Espeland, M. 2019. A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae). ZooKeys 821: 85-152. DOI: 10.3897/zookeys.821.31782 (31 Jan 2019).

2018. Espeland, M., Breinholt, J., Barbosa, E., Casagrande, M. M., Huertas, B., Lamas, L., Marín, M. A., Mielke, O. H. H., Miller, J. Y., Nakahara, S., Tan, D., Warren, A. D., Zacca, T., Kawahara, A., Freitas, A. V. L., Willmott, K. R. Four hundred shades of brown: Higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data. Molecular Phylogenetics and Evolution 131: 116-124, DOI: 10.1016/j.ympev.2018.10.039 (online 10 Nov 2018).

2018. Checa, M. F., Donoso, D. A., Rodríguez, J., Levy, E., Warren, A., Willmott, K. R. Combining sampling techniques aids monitoring of tropical butterflies. Insect Conservation and Diversity DOI: 10.1111/icad.12328 (online 29 Oct 2018).

2018. Dufour, P. C., Willmott, K. R., Padrón, P. S., Xing, S., Bonebrake, T. C., Scheffers, B. R. Divergent melanism strategies in Andean butterfly communities structure diversity patterns and climate responses. Journal of Biogeography 45: 2471-2482. DOI: 10.1111/jbi.13433 (online 4 October 2018).

2018. Rosser, N., Freitas, A. V. L., Huertas, B., Joron, M., Lamas, G., Mérot, C., Simpson, F., Willmott, K. R., Mallet, J., Dasmahapatra, K. A. Cryptic speciation associated with geographic and ecological divergence in two Amazonian Heliconius butterflies. Zoological Journal of the Linnean Society XX: 1-17 DOI: 10.1093/zoolinnean/zly046 (online 4 August 2018).

2018. Dias, F. M. S., Janzen, D., Hallwachs, W., Chacón, I., Willmott, K. R., Ortiz-Acevedo, E., Mielke, O. H. H., Casagrande, M. M. DNA Barcodes uncover hidden taxonomic diversity behind the variable wing patterns in the Neotropical butterfly genus Zaretis (Lepidoptera: Nymphalidae: Charaxinae). Zoological Journal of the Linnean Society XX: 1-61. DOI: 10.1093/zoolinnean/zly036 (online 1 August 2018).

2018. Nakahara, S., Willmott, K. R., Mielke, O. H. H., Schwartz, J., Zacca, T., Espeland, M., Lamas, G. Seven new taxa from the butterfly subtribe Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) with revisional notes on Harjesia Forster, 1964 and Pseudeuptychia Forster, 1964. Insecta Mundi 0639: 1-38 (online 27 July 2018).

2018. Willmott, K. R., Lamas, G., Radford, J., Marín, M. A., Nakahara, S., Espeland, M., Xiao, L., Hall, J. P. W. A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru. Tropical Lepidoptera Research 28(1): 39-45 (online 13 July 2018).

2018. Zacca, T., Casagrande, M. M., Mielke O. H. H., Huertas, B., Barbosa E. P., Freitas, A. V. L., Magaldi, L. M., Espeland, M., Nakahara, S., Willmott, K. R. Systematics of the butterfly genus Cissia Doubleday, 1848 (Lepidoptera: Nymphalidae, Satyrinae) using an integrative approach. Arthropod Systematics and Phylogeny, 76(2): 349-376 (June).

2018. Benmesbah, M., Zacca, T., Casagrande, M. M., Mielke, O. H. H., Lamas, G., Willmott, K. R. Taxonomic notes on Papilio ocypete Fabricius, 1776 and Papilio helle Cramer, 1779 with description of two new similar species from South America (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4425(1): 115-145 (May).

2018. Barbosa, E. P., Siewert, R. R., Mielke, O. H. H., Lamas, G., Willmott, K. R., Freitas, A. V. L. Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae). Zootaxa, 4422(4): 537-557, DOI: 10.11646/zootaxa.4422.4.5 (online 25 May 2018).

2018: Freitas, A. V. L., Barbosa, E., Zacca, T., Marín, M., Beirão, M., Silva, A., Casagrande, M. M., Espeland, M., Willmott, K. R. 2018. Before it is too late: description of a new genus and species of butterfly from a highly threatened Brazilian biome. Revista Brasileira de Entomologia 62: 148-158, DOI: 10.1016/j.rbe.2018.02.003 (online 9 May).

2018. Finkbeiner, S. D., Salazar-Carrión, P., Nogales, S., Rush, C., Briscoe, A. D., Hill, R., Kronforst, M., Willmott, K. R., Mullen, S. P. Frequency-dependence shapes the adaptive landscape of imperfect Batesian mimicry. Proceedings of the Royal Society B 285: 20172786 (online 4 April 2018).

2018. Chazot, N., De Silva, D. L., Willmott, K. R., Freitas, A. V. L., Lamas, G., Mallet, J., Giraldo, C. E., Uribe, S., Elias, M. Contrasting patterns of Andean diversification among three diverse clades of Neotropical clearwing butterflies. Ecology and Evolution, 2017; 1-18, DOI: 10.1002/ece3.3622 (online 25 March 2018).

2018. Espeland, M., Breinholt, J., Willmott, K. R., Warren, A. D., Vila, R., Toussaint, E. F. A., Maunsell, S. C., Aduse-Poku, K., Talavera, G., Eastwood, R., Jarzyna, M. A., Ries, L., Guralnick, R., Lohman, D. J., Pierce, N. E., Kawahara, A. Comprehensive higher-level phylogeny of butterflies (Papilionoidea) inferred from genomic data. Current Biology, 28: 770-778, DOI: 10.1016/j.cub.2018.01.061 (5 March).

2017. Ortiz-Acevedo, E., Bonfantti, D., Casagrande, M., Mielke, O. H. H., Espeland, M., Willmott, K. R. Using molecules and morphology to unravel the systematics of Neotropical preponine butterflies (Lepidoptera: Charaxinae: Preponini). Insect Systematics and Diversity, 1: 48-56, DOI: 10.1093/isd/ixx002 (online 25 October).

2017. Nakahara, S., Zacca, T., Huertas, B., Neild, A. F. E., Hall, J. P. W., Lamas, G., Holian, L. A., Espeland, M., Willmott, K. R. Remarkable sexual dimorphism, rarity and cryptic species: a revision of the ‘aegrota species group’ of the Neotropical butterfly genus Caeruleuptychia Forster, 1964 with the description of three new species (Lepidoptera, Nymphalidae, Satyrinae). Insect Systematics and Evolution, 1-53, DOI: 10.1163/1876312X-00002167 (online 31 July).

2017. Willmott, K. R., Lamas, G., Hall, J. P. W. Notes on the taxonomy of Actinote intensa Jordan (Lepidoptera: Nymphalidae: Heliconiinae) and the description of a new sibling species from eastern Ecuador. Tropical Lepidoptera Research, 27(1): 6-15 (June).

2017. Willmott, K. R., Simon, M., Ortiz-Acevedo, E., Hall, J. P. W. First record of the enigmatic tribe Anaeomorphini (Lepidoptera, Nymphalidae, Charaxinae) outside of the Amazon basin: a new species of Anaeomorpha Rothschild, 1894, from the Chocó region of western Ecuador. Insecta Mundi, 0548: 1-10 (May).

2017. Willmott, K. R., Robinson Willmott, J. C., Elias, M., Jiggins, C. Maintaining mimicry diversity: optimal warning color patterns differ among microhabitats in Amazonian clearwing butterflies. Proceedings of the Royal Society B, 284: 20170744. DOI: 10.1098/rspb.2017.0744 (May).

2017. De Silva, D. L., Mota, L., Chazot, N., Mallarino, R., Silva-Brandão, K. L., Gómez P., L. M., Freitas, A. V. L., Lamas, G., Joron, M., Mallet, J., Giraldo S., C. E., Uribe S., S. I., Särkinen, T., Knapp, S., Jiggins, C. D., Willmott, K. R., Elias, M. 2017. North Andean origin and diversification of the largest ithomiine butterfly genus. Scientific Reports, 7: 45966, DOI: 10.1038/srep45966 (April).

2016. Nakahara, S., Tan, D., Lamas, G., Parus, A., Willmott, K. R. 2016. A distinctive new species of Hermeuptychia Forster, 1964 from the eastern tropical Andes (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research, 26: 77-84 (December).

2016. Freitas, A. V. L., Magaldi, L. M., Willmott, K. R. A new subspecies of Oleria gunilla (Nymphalidae: Danainae) from north Mato Grosso, Brazil. Journal of the Lepidopterists’ Society, 70(4): 295-301.

2016. Nakahara, S., Vega, G., Willmott, K. R. Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes. Zootaxa, 4184: 358-366. DOI: 10.11646/zootaxa.4184.2.7 (November).

2016. Huertas, B., Lamas, G., Fagua, G., Mallet, J., Nakahara, S., Willmott, K. R. A remarkable new butterfly species from western Amazonia (Lepidoptera: Nymphalidae: Satyrinae). Conservación Colombiana, 24: 1-5. (October).

2016. Chazot, N., Willmott, K. R., Condamine, F., de Silva, D. L., Freitas, A. V. L., Lamas, G., Morlon, H., Giraldo, C. E., Jiggins, C. D., Joron, M., Mallet, J., Uribe, S., Elias, M. Into the Andes: multiple independent colonizations drive montane diversity in the Neotropical clearwing butterflies Godyridina. Molecular Ecology DOI: 10.1111/mec.13773 (October).

2016. Willmott, K. R., Mielke, O. H. H., Freitas, A. V. L. A new subspecies and validation of the enigmatic species Adelpha herbita Weymer endemic to the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research, 26: 32-37 (June).

2016. Nakahara, S., Barbosa, E. P., Marín, M. A., Freitas, A. V. L., Pomerantz, T., Willmott, K. R. Graphita gen. nov., a new genus for Neonympha griphe C. Felder & R. Felder, 1867 (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology, 17 pp, doi: 10.1007/s13744-016-0414-x (June).

2016. Freitas, A. V. L., Barbosa, E., Willmott, K. R., Wahlberg, N., Lamas, G. ‘Species’ from three different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileira de Entomologia, doi:10.1016/j.rbe.2016.01.004 (March).

2015. Nakahara, S., Hall, J. P. W., Lamas, G., Willmott, K. R. Seven new species and one new subspecies of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the tropical Andes. Tropical Lepidoptera Research 25(2): 63-79.

2015. Garzón-Orduña, I. J., Silva-Brandão, K. L., Willmott, K. R., Freitas, A. V. L., Brower, A. V. Z. Incompatible ages for clearwing butterflies based on alternative secondary calibrations. Systematic Biology DOI:10.1093/sysbio/syv032.

2015. De-Silva, D. L., Elias, M., Willmott, K. R., Mallet, J., Day, J. J. Diversification of clearwing butterflies with the rise of the Andes. Journal of Biogeography (online September).

2015. Willmott, K. R. A new subspecies of Cithaerias pyropina (Salvin & Godman, 1868) from the Cordillera de Cutucú in southeastern Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera Research, 25(1): 1-4.

2015. Ebel, E. R., DaCosta, J. M., Sorenson, M. D., Hill, R. I., Briscoe, A. D., Willmott, K. R., Mullen, S. P. Rapid diversification associated with ecological specialization in Neotropical Adelpha butterflies. Molecular Ecology, 24: 2392-2405.

2014. Warren, A. D., Tan, D., Willmott, K. R., Grishin, N. V. Refining the diagnostic characters and distribution of Hermeuptychia intricata (Nymphalidae: Satyrinae: Satyrini). Tropical Lepidoptera Research, 24: 44-51. Download reprint [PDF]

2014. Brower, A. V. Z., Willmott, K. R., Silva-Brandão, K. L., Garzón-Orduña, I. J., Freitas, A. V. L. Phylogenetic relationships of ithomiine butterflies (Lepidoptera: Nymphalidae: Danainae) as implied by combined morphological and molecular data. Systematics and Biodiversity, DOI: 10.1080/14772000.2014.899650, 1-15.

2014. Checa, M. F., Rodríguez, J., Willmott, K. R., Liger, B. Microhabitat significantly affects the composition, abundance and phenology of butterfly assemblages in an Ecuadorian dry forest. Florida Entomologist, 97: 1-13.

2013. Chazot, N., Willmott, K. R., Paola G., S. E., Toporov, A., Hill, R., Jiggins, C., Elias, M. Mutualistic mimicry and filtering by altitude shape the structure of Andean butterfly communities. American Naturalist, doi: 10.1086/674100.

2013. Willmott, K. R., Hall, J. P. W. A new species and two new subspecies of Adelpha Hübner, [1819] from the tropical Andes (Lepidoptera: Limenitidinae).Journal of the Lepidopterists’ Society, 67: 241-252. Download reprint [PDF]

2013. Radford, J., Willmott, K. R. A distinctive new subspecies of Catasticta poujadei from the Cordillera del Cóndor in eastern Ecuador (Lepidoptera: Pieridae: Pierinae). Tropical Lepidoptera Research, 23: 86-91. Download reprint [PDF]

2013. Pyrcz, T. W., Willmott, K. R., Garlacz, R., Boyer, P. Latitudinal gradient and spatial covariance in species richness of tropical Lepidoptera in the Andes.Insect Diversity and Conservation, Online version: DOI: 10.1111/icad.12058.

2013. Ortíz-Acevedo, E., Willmott, K. R. Molecular systematics of the butterfly tribe Preponini (Nymphalidae: Charaxinae). Systematic Entomology. Online version: DOI: 10.1111/syen.12008.

2013. Giraldo, C. E., Willmott, K. R., Vila, R. Uribe, S. I. Ithomiini butterflies (Lepidoptera: Nymphalidae) of Antioquia, Colombia. Neotropical Entomology.Online version: DOI: 10.1007/s13744-012-0102-4.

2012. Matos-Maraví, P. F., Peña, C., Willmott, K. R., Freitas, A. V. L., Wahlberg, N. Systematics and evolutionary history of butterflies in the “Taygetis clade” (Nymphalidae: Satyrinae: Euptychiina): towards a better understanding of Neotropical biogeography. Molecular Phylogenetics and Evolution, 66: 54-68.Online version: DOI: 10.1016/j.ympev.2012.09.005

2012. Hill, R. I., Elias, M., Dasmahapatra, K. K., Jiggins, C. D., Koong, V., Willmott, K. R., Mallet, J. Ecologically relevant cryptic species in the highly polymorphic Amazonian butterfly Mechanitis mazaeus s.l. (Lepidoptera: Nymphalidae; Ithomiini). Biological Journal of the Linnean Society, 106: 540-560. Online version: DOI: 10.1111/j.1095-8312.2012.01874.x

2012. Rosser, N., Phillimore, A.B., Huertas, B., Willmott, K.R., Mallet, J. Testing historical explanations for gradients in species richness in heliconiine butterflies of tropical America. Biological Journal of the Linnean Society, 105: 479-497. Online version: DOI: 10.1111/j.1095-8312.2011.01814.x.

2011. Willmott, K. R., Elias, M., Sourakov, A. Two possible caterpillar mimicry complexes in neotropical danaine butterflies (Nymphalidae, Danainae: Danaini, Ithomiini). Annals of the Entomological Society of America, 104: 1108-1118. Abstract.

2011. Gompert, Z., Willmott, K. R., Elias, M. Heterogeneity in predator micro-habitat use and the maintenance of Müllerian mimicry diversity. Journal of Theoretical Biology, 281: 39-46. Online version.

2011. Bolaños M., I. A., Zambrano G., G., Willmott, K. R. Descripción de los estados inmaduros de Pteronymia zerlina zerlina, P. zerlina machay, P. medellina y P. veia n. ssp. (Lepidoptera: Nymphalidae: Ithomiini) de Colombia y del Ecuador. Tropical Lepidoptera Research, 21: 27-33. Download reprint [PDF]

2011. Pyrcz, T., Greeney, H. F., Willmott, K. R., Wojtusiak, J. A taxonomic revision of the genus Daedalma Hewitson with the descriptions of twenty new taxa and the immature stages of two species (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 2898: 1-68. Online version. Abstract

2010. Mullen, S. P., Savage, W., Willmott, K. R. Rapid diversification and not clade age explains high diversity in neotropical Adelpha butterflies. Proceedings of the Royal Society of London B doi:10.1098/rspb.2010.2140 Online version

2010. Hall, J. P. W., Willmott, K. R. Description of a new Lucillella species (Riodinidae: Symmachiini) discovered in the eastern Andes of Ecuador using the single rope canopy access technique. Journal of the Lepidopterists’ Society, 64: 139-146. DOI 10.18473/lepi.v64i3.a2

2010. Willmott, K. R., Hall, J. P. W. A new species of Dynamine Hübner, [1819] from northwestern Ecuador (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera Research, 20: 23-27. Download reprint [PDF]

2010. de-Silva, D. L., Day, J. J., Elias, M., Willmott, K. R., Whinnett, A., Mallet, J. Molecular phylogenetics of the neotropical butterfly subtribe Oleriina (Nymphalidae: Danainae: Ithomiini). Molecular Phylogenetics and Evolution, 55:1032-1041. Download reprint [PDF]

2009. Janzen, D. J., Hallwachs, W., + 39 authors, Willmott, K. R., + 3 authors. Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity. Molecular Ecology Resources, 9(Suppl. 1), 1-26. Online version

2009. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Phylogenetic community ecology needs to take positive interactions into account: Insights from colourful butterflies. Communicative & Integrative Biology 2(2): 113-116 [Article addendum]. Download reprint [PDF]

2009. Hall, J. P. W., and K. R. Willmott. Two new species of Euselasia (Riodinidae: Euselasiinae) from western Ecuador. Tropical Lepidoptera Research, 19(1): 52-55. Download reprint [PDF]

2009. Elias, M., Joron, M., Willmott, K. R., Kaiser, V., Silva-Brandão, K. L., Freitas, A. V. L., Arias, C. F., Gomez Piñeres, L. M., Uribe, S., Jiggins, C. Out of the Andes: patterns of diversification in clearwing butterflies. Molecular Ecology, 18: 1716-1729. Download page proofs [PDF]

2009. Willmott, K. R., Freitas, A. V. L., Hall, J. P. W., Silva-Brandão, K. L., Paluch, M. A new species of Actinote Hübner from the eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56. Download reprint [PDF] – NOTE: figures in published version are monochrome. DOI 10.4289/0013-8797-111.1.47

2008. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Mutualistic interations drive ecological niche convergence in a diverse butterfly community. PLoS Biology, 6(12):e300. Definitive online version or Download reprint [PDF], Supporting data S1 [PDF], Supporting data S2 [PDF]

2008. Willmott, K. R., Vitale, F. Taxonomic notes on Napeogenes (Lepidoptera: Ithomiinae) from Ecuador and Colombia, with the description of ten new subspecies. Genus, 19(3): 585-609. Download reprint [PDF]

2008. Willmott, K. R., Lamas, G. A revision of the genus Megoleria (Lepidoptera, Nymphalidae, Ithomiinae). Tropical Lepidoptera Research, 18(1): 46-59. Download reprint [PDF]

2007. Elias, M., Hill, R., Dasmahapatra, K., Willmott, K. R., Brower, A., Mallet, J., and C. Jiggins. Limited performance of DNA barcoding in a diverse community of tropical butterflies. Proceedings of the Royal Society of London B, 274: 2881-2889. Download reprint [PDF] and supplementary documentation

2007. Brehm, G., Hartmann, T., and K. R. Willmott. Pyrrolizidine alkaloids and pharmacophagous Lepidoptera visitors of Prestonia amabilis (Apocynaceae) in a montane rain forest in Ecuador. Annals of the Missouri Botanical Garden, 94(2): 463-473. Download reprint [PDF]

2007. Hall, J. P. W., and K. R. Willmott. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 1-5. Download reprint [PDF]

2007. Willmott, K. R., and G. Lamas. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America, 100: 449-469. Download reprint [PDF]

2006. Pyrcz, T. W., Willmott, K. R., Hall, J. P. W., and A. L. Viloria. A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39: 37-79 (2000). Download reprint [PDF]

2006. Willmott, K. R., and A. V. L. Freitas. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics, 22: 297-368. Online version, Download PAGE PROOFS [PDF] or e-mail authors for reprint

2006. Jiggins, C. D., Mallarino, R., Willmott, K. R., and E. Bermingham. The phylogenetic pattern of speciation and wing pattern change in neotropicalIthomia butterflies (Lepidoptera: Nymphalidae). Evolution, 60: 1454-1466. Download reprint [PDF]

2006. Willmott, K. R., and G. Lamas. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology, 31(3): 419-468. Download reprint [PDF]

2006. Brower, A. V. Z., Freitas, A. V. L., Lee, M.-M., Silva Brandão, K. L., Whinnett, A., and K. R. Willmott. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology, 31: 288-301. Download reprint [PDF]

2005. Whinnett, A., Willmott, K. R., Brower, A.V.Z., Simpson, F., Zimmermann, M., Lamas, G., and J. Mallet. Mitochondrial DNA provides an insight into the mechanisms driving diversification in the ithomiine butterfly Hyposcada anchiala (Lepidoptera: Nymphalidae, Ithomiinae). European Journal of Entomology, 102: 633-639. Download reprint [PDF]

2005. Whinnett, A., Zimmermann, M., Willmott, K. R., Herrera, N., Mallarino, R., Simpson, F., Joron, M., Lamas, G., and J. Mallet. Strikingly variable divergence times inferred across an Amazonian butterfly ‘suture zone’. Proceedings of the Royal Society of London B, 272: 2525-2533. Download reprint [PDF]

2005. Whinnett, A., Brower, A. V. Z., Lee, M.-M., Willmott, K. R., and J. Mallet. The phylogenetic utility of tektin, a novel region for inferring systematic relationships amongst Lepidoptera. Annals of the Entomological Society of America, 98: 873-886. Download reprint [PDF]

2005. Mallarino, R., Bermingham, E., Willmott, K. R., Whinnett, A., and C. D. Jiggins. Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes. Molecular Phylogenetics and Evolution, 34(3): 625-644. Download reprint [PDF]

2005. Hall, J. P. W., K. R. Willmott and R. C. Busby. Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa, 797: 1-20. Download reprint [PDF]

2004. Balcázar-Lara, M., and K. R. Willmott. A new subspecies of Adelpha erymanthis from Mexico, with a key to identification of similar taxa (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera, 12(1-2)(“2001”): 25-38. Download reprint [PDF]

2004. Willmott, K. R., and J. Mallet. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Suppl.) 271: S266-S269. Download reprint [PDF]

2004. Willmott, K. R., and Lamas, G. A new species of Tithorea (Lepidoptera: Nymphalidae: Ithomiinae) from the Chocó region of northwestern South America and Panama. J. New York Ent. Soc., 112(1): 1-17. Download reprint [PDF]

2004. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on the genus Zaretis, with the description of a new species (Lepidoptera: Nymphalidae: Charaxinae). Tropical Lepidoptera, 12(1-2): 25-38. Download reprint [PDF]

2003. Mallet, J. L. B., and K. R. Willmott. Taxonomy: renaissance or Tower of Babel? Trends in Ecology and Evolution, 18(2): 57-59. Online version

2003. Willmott, K. R. Cladistic analysis of the neotropical butterfly genus Adelpha (Nymphalidae: Limenitidini), with comments on the subtribal classification of the tribe Limenitidini. Systematic Entomology, 28: 1-43. Download reprint [PDF]

2002. Robinson-Dean, J. C., K. R. Willmott, M. J. Catterall, D. J. Kelly, A. Whittington, B. Phalan, N. M. Marples, and D. R. S. Boeadi. A new subspecies of Red-backed Thrush Zoothera erythronota kabaena subsp. nov. (Muscicapidae: Turdidae), from Kabaena island, Indonesia. Forktail, 18: 1-10.

2001. Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology, 26: 369-399. Download reprint [PDF]

2001. Willmott, K. R., L. M. Constantino, and J. P. W. Hall. A review of Colobura (Lepidoptera: Nymphalidae) with comments on larval and adult ecology and description of a sibling species. Annals of the Entomological Society of America, 94(2): 185-196. Download reprint [PDF]

2000. Hall, J. P. W., and K. R. Willmott. Patterns of feeding behaviour in adult male riodinid butterflies and their relationship to morphology and ecology. Biol. J. Linn. Soc., 69: 1-23. Download reprint [PDF]

1999. Pyrcz, T., K. R. Willmott, and J. P. W. Hall. Contribution to the knowledge of Ecuadorian Pronophilini. Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus, 10(3): 497-522. Download reprint [PDF – 1] [PDF – 2]

1999. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on Ecuadorian Adelpha with the description of two new species and seven new subspecies (Lepidoptera: Nymphalidae: Limenitidinae). Tropical Lepidoptera, 10(1): 1-17. Download reprint [PDF – 1] [PDF – 2]

1998. Hall, J. P. W., and K. R. Willmott. Four new riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea, 98: 325-334. Download reprint [PDF]

1998. Hall, J. P. W., and K. R. Willmott. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 27-35. Download reprint [PDF]

1998. Hall, J. P. W., and K. R. Willmott. Three new species of Riodinini from the cloud forests of Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 22-26. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 63-67. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana Part II, with the description of a new species. Tropical Lepidoptera, 7(1): 71-80. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea, 96: 637-660. Download reprint [PDF]

1996. Torres, R., J. P. W. Hall, K. R. Willmott, and K. Johnson. A new genus of brilliantly colored elfin butterflies from the high Andes (Lepidoptera: Lycaenidae). Tropical Lepidoptera, 7(1): 81-86. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 110-112. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae).Tropical Lepidoptera, 6(2): 131-135. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 136-143. Download reprint [PDF]

1995. Willmott, K. R., and J. P. W. Hall. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera, 6(2): 103-105. Download reprint [PDF]

1995. Willmott, K. R., and J. P. W. Hall. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 6(2): 106-109. Download reprint [PDF]

1994. Willmott, K. R., and J. P. W. Hall. Four new species of Riodinidae (Lepidoptera) from Western Ecuador. Tropical Lepidoptera, 5(2): 87-91. Download reprint [PDF]

1993. Hall, J. P. W., and K. R. Willmott. Temporal and elevational additions of distribution for Costa Rican butterflies. Tropical Lepidoptera, 4(2): 139-141. Online version

Books

2003. Willmott, K. R. The Genus Adelpha: its Systematics, Biology and Biogeography (Lepidoptera: Nymphalidae: Limenitidini). Gainesville, Scientific Publishers. viii + 322 pp. Download below:

NOTE: File sizes are quite large (2-10 MB). If you can afford it, please buy a copy of the book from Scientific Publishers by contacting Dr John Heppner at jheppner@flmnh.ufl.edu

Complete book (c. 45 Mb – compiled text sections and figures are available individually below)

Individual text sections

Figures

 

Book chapters

2014. Checa, M. F., Willmott, K. R. Diversidad e importancia ecológica de mariposas del Ecuador. In: García, M., Parra, D. (Eds.) Ecuador: país de la Biodiversidad. Fundación Botánica de Los Andes/Ministerio del Ambiente/Fundación Ecofondo. Quito, Ecuador.

2006. Gareca, Y., Forno, E., Pyrcz, T., Willmott, K. R., Reichle, S. Lista preliminar de mariposas diurnas de Bolivia. In: Gareca, Y., and S. Reichle (eds.),Mariposas Diurnas de Bolivia, pp. 1-66. Tarija, Bolivia, PROMETA.

2004. Willmott, K.R. Brush-footed butterflies (Lepidoptera: Nymphalidae). In: Capinera, J. L. (Ed.), Encyclopedia of Entomology, Volume I A-E, pp. 379-384. Dordrecht / Boston / London, Kluwer Academic Publishers. Download reprint [PDF]

Popular publications

2004. Willmott, K. R., and J. P. W. Hall. Common Butterflies of Yuturi and Yarina lodge. Yuturi Lodge, Ecuador. 2 pp.

2001. Hall, J. P. W., and K. R. Willmott. Ecuador. Out on a limb. National Geographic Magazine (Update from the field), 199(2): 86. Download reprint [PDF]

2001. Willmott, K. R., and J. P. W. Hall. Winged gems of Ecuador. Biodiversity paradise. National Geographic Magazine (Geographica) 199(5): xxviii. Download reprint [PDF]

Letters and reviews

2015. Willmott, K. R. Butterflies. By Orenstein, R. I., Marent, T. (2015). Tropical Lepidoptera Research 25(2): 104-105. [book review]

2014. Warren, A. D., Willmott, K. R., Grishin, N. V. Subtle satyrs: differentiation and distribution of the newly described Hermeuptychia intricata in the southeastern United States (Lepidoptera: Nymphalidae: Satyrinae). News of the Lepidopterists’ Society 56(2): 83-85.

2012. Willmott, K. R. Lepidoptera. Borboletas e Mariposas do Brasil. [Lepidoptera. Butterflies and Moths of Brazil.] By de Almeida, A. C., Freitas, A. V. L. (2012). Tropical Lepidoptera Research 22(2): 120. [book review]

2009. Willmott, K. R. Mariposas de Canandé. Sus Amenazas, Potencial y Futuro. [The Butterflies of Canandé. Their Threats, Potential and Future.] By Checa V., M. F. (2009). Tropical Lepidoptera Research 19(1): 60. [book review]

2008. Willmott, K. R. Catalogue of the Hostplants of the Neotropical Butterflies/ Catálogo de las Plantas Huésped de las Mariposas Neotropicales. By Beccaloni, G. W., Viloria, A. L., Hall, S. K. & Robinson, G. S. (2008). Tropical Lepidoptera Research 18(2): 124-125. [book review]

1998. Willmott, K. R. A fair review? News of the Lepidopterist’s Society, 40(4): 92-93.

1997. Hall, J. P. W., and K. R. Willmott. The Butterflies of Venezuela. Part I. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae). A Comprehensive Guide to the Identification of Adult Nymphalidae, Ppailionidae and Pieridae. By Andrew F. E. Neild. Tropical Lepidoptera, 8(1): 46. [book review]